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Acetyl-DL-carnitine levels were significantly higher in untreated R cells than WT cells (p-value = 5.8 10 12 ), whereas hexanoyl-L-carnitine was significantly higher in untreated WT cells than R (p-value = 6.6 10 9 )

A weak interaction, which could be lost during protein purification, would explain the lack of EtfD-mediated pumping activity by IMVs that are not supplemented with purified ETF